Page 6 - LN-CH-2[2]
P. 6

Double fertilization:


                             After entering one of the synergids, the pollen tube releases two male gametes into the cytoplasm
                              of the synergids.
                             Syngamy: one of the male gamete fused with egg cell, to form a diploid zygote.
                             Two polar nuclei of central cell fused to form a diploid secondary nucleus.
                             Triple  fusion:  The  second  male  gamete  fused  with  the  secondary  nucleus  to  form  a
                              triploid primary endosperm nucleus.
                             Since two type of fusion, syngamy and triple fusion take place in the embryo sac the phenomenon
                              is termed asdouble fertilization.
                             The  central  cell  after  triple  fusion  becomes primary  endosperm  cell and  developed  into
                              the endosperm.
                             The zygote developed into an embryo.


               POST- FERTILIZATION : STRUCTURE AND EVENTS

               Events of endosperm and embryo development, maturation of ovule into seed and ovary into fruit, are collectively
               termed aspost-fertilization events.

               Endosperm:


                             Development of endosperm takes place before the embryo development.
                             Primary endosperm cell divides repeatedly to form a triploid endosperm.
                             Cells are filled with reserve food material and are used for the nutrition of the developing embryo.
                             PEN undergoes successive nuclear division to give rise to free nuclei. This is called free-nuclear
                              endosperm.
                             Subsequently cell wall formation takes place and become cellular endosperm.
                             The coconut water is free nuclear endosperm and the white kernel is the cellular endosperm.
                             Endosperm may be consumed completely during embryo developed or it may be consumed during
                              germination of seed.

               Embryo:

                             Zygote formed and placed at the micropylar end of the embryo sac.
                             Zygote starts its development only after some amount of endosperm formed.
                             Embryo development takes place in following stages:
                                 o   Proembryo
                                 o   Globular stage
                                 o   Heart shaped
                                 o   Matured embryo.

               Dicot embryo:


                             A typical dicotyledonous embryo consists of an embryonal axis and two cotyledons.
                             Embryonal axis above the cotyledon is the epicotyls.
                             Terminal part of the epicotyls is the plumule (gives rise to the shoot).
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